Revolution water-cooled engine Revolution, 2002-present, 69 cubic inches (1.13 L), 60 degree V-twin, Liquid cooled. Used in VRSC (V-Twin Racing Street Custom) models. At first it was used in a single model called the V-Rod; it now comes in four consumer models: The V-Rod, the "roadster-styled" Street Rod, the long and low Night Rod, and the VRSCSE² Screamin’ Eagle V-Rod. There is also a non-street fifth variant: the VRXSE SE Destroyer. The Screamin’ Eagle V-Rod Destroyer is a 165+hp, professional-level drag-racing motorcycle designed exclusively for competition by Harley-Davidson Custom Vehicle Operations; it comes stock with larger 79 cu.in. engine, air shifter and wheelie bar, among other features. The Destroyer is a turn-key motorcycle tuned to make sub 10-second quarter-mile runs in the hands of a pro-level rider.

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The company considers 1903 to be its year of founding, though the Harley-Davidson enterprise could be considered to have started in 1901 when William S. Harley, age 21, drew up plans for a small engine that displaced 7.07 cubic inches (116cc) and had four-inch flywheels. The engine was designed for use in a regular pedal-bicycle frame. Over the next two years Harley and his boyhood pal Arthur Davidson labored on their little motor-bicycle using the northside machine shop of their friend Henry Melk. It was finished in 1903 with the help of Arthur's brother, Walter Davidson. Upon completion the boys found their little "power-cycle" unable to conquer Milwaukee's modest hills without pedal assistance. Will Harley and the Davidsons quickly wrote off their first motor-bicycle as a valuable learning experiment.

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